Tropical Grasslands-Forrajes Tropicales (2018) Vol. 6(1):34–41 34

DOI: 10.17138/TGFT(6)34-41

Research Paper

Evaluation of growth parameters and forage yield of Sugar Graze

and Jumbo Plus sorghum hybrids under three different spacings

during the maha season in the dry zone of Sri Lanka

Efecto de distancia de siembra en el desarrollo y rendimiento de dos

híbridos de sorgo forrajero (Sugar Graze y Jumbo Plus) durante la

temporada de maha en la zona seca de Sri Lanka

HAJAROOBA GNANAGOBAL AND JEYALINGAWATHANI SINNIAH

Faculty of Agriculture, University of Jaffna, Ariviyal Nagar, Kilinochchi, Sri Lanka. www.agri.jfn.ac.lk

Abstract

A field experiment to evaluate the growth parameters and fodder yields of Sugar Graze and Jumbo Plus under occasional irrigation was conducted at 3 different plant spacings (30 × 15, 30 × 45 and 30 × 60 cm) on a red-yellow latosol in the dry zone of Sri Lanka from August 2015 to January 2016. The design was a randomized block with 3 replications. Initial harvesting of fodder was done 60 days after planting and 2 ratoon yields were assessed at successive 60-day intervals.

Plant spacing was inversely related (P<0.05) to dry matter (DM) yield with the narrowest spacing (30 × 15 cm) producing yields of 14.1 t DM/ha for Sugar Graze and 12.6 t DM/ha for Jumbo Plus at the initial harvest. Plant spacing also influenced leaf area, stem girth, root length and plant height in the initial harvest. Sugar Graze produced higher yields than Jumbo Plus at the initial and second ratoon harvests. Yields from ratoon crops were about 30% of those for the initial harvest. Further studies are needed to determine how these findings apply under the low-rainfall conditions of the yala season, and chemical analyses and animal feeding studies would provide valuable information on the nutritional value of the different forages.

Keywords: Dry matter yield, forage sorghum, ratoon crop, red yellow latosol, row width.

Resumen

En un latosol rojo-amarillo de la zona seca al norte de Sri Lanka entre agosto de 2015 y enero de 2016 se evaluaron algunas características de crecimiento y los rendimientos de forraje de los cultivares Sugar Graze y Jumbo Plus bajo riego ocasional usando 3 distancias de siembra (30 × 15, 30 × 45 y 30 × 60 cm). Los tratamientos se dispusieron en un diseño de bloque al azar con 3 repeticiones. La primera cosecha de forraje se realizó 60 días después de la siembra, seguida por 2 cosechas de rebrote a un intervalo de 60 días cada una. Los resultados mostraron que la distancia de siembra se relacionó de manera inversa (P<0.05) con el rendimiento de materia seca (MS), siendo este más alto (14.1 t MS/ha) en la distancia 30 × 15 cm en la primera cosecha para el cv. Sugar Graze en comparación con el cv. Jumbo Plus (12.6 t MS/ha). La distancia de siembra también influyó en el área foliar, el grosor del tallo, la longitud de las raíces y la altura de la planta en la primera cosecha. Sugar Graze produjo mayores rendimientos que Jumbo Plus en la primera cosecha y en la segunda cosecha de rebrote. Los rendimientos en las dos cosechas de rebrote fueron de alrededor del 30% de la primera cosecha. Se requieren estudios adicionales para determinar cómo se comparan estos resultados con los que se puedan obtener en época seca (temporada yala). Además, análisis químicos y estudios nutricionales con animales proporcionarían información valiosa sobre el valor nutritivo de los diferentes forrajes.

Palabras clave : Distancia entre surcos, latosol amarillo-rojo, rebrote, rendimiento de materia seca.

___________

Correspondence: Dr. (Ms.) Jeyalingawathani Sinniah, Department of

Animal Science, Faculty of Agriculture, University of Jaffna, Ariviyal

Nagar, Kilinochchi, 44000, Sri Lanka.

E-mail: jeyalingawathani@gmail.com

Tropical Grasslands-Forrajes Tropicales (ISSN: 2346-3775)

Yield parameters of forage sorghum hybrids 35

Introduction

cultivars in the dry zone of Sri Lanka are limited. There is

a need to assess the growth of these cultivars and how

High performance of farm animals, especially dairy cows,

varying the plant spacing affects both yield and quality of

depends on the availability of adequate amounts of quality

forage so that the growing demand for forage by livestock

fodder and in developing countries, inadequacy of quality

can be met.

forage is the critical limitation to profitable animal

The present study was designed to determine the crop

production (Sarwar et al. 2002). Among the many options

morphology, growth parameters and forage yield of Sugar

for overcoming the shortage of forage, the introduction of

Graze and Jumbo Plus under irrigation in the dry zone of

high-yielding crop varieties ranks highly (Bilal et al.

Sri Lanka under 3 different plant spacings (30 × 15, 30 ×

2001). However, in many developing countries, because

45 and 30 × 60 cm).

of the ever-growing need for food for humans, only

limited cultivated land can be allocated to produce fodder

Materials and Methods

for livestock. Douglas (1980) recommended annual

summer crops such as forage sorghum hybrids ( Sorghum

This experiment was carried out at the livestock farm,

spp.) for use as alternative forage crops in drier areas in

Department of Animal Science, Faculty of Agriculture,

order to bridge the feed shortage gap.

University of Jaffna, Ariviyal Nagar, Kilinochchi (Figure

Sugar Graze, a sweet sorghum × sweet sorghum

1), from August 2015 to January 2016. These months fall

hybrid , is a popular forage source among the livestock

into Sri Lanka’s maha season, i.e. the period September –

farmers of Sri Lanka and Jumbo Plus, a sweet sorghum ×

February which experiences rainfall through the

Sudan grass hybrid, is still in the initial stages of

Northeast monsoon.

introduction. Sugar Graze is a late-flowering cultivar with

high yields, a crude protein (CP) concentration of 12‒

18% and a high sugar content that boosts feed quality and

palatability, resulting in minimal feed wastage. In

addition, the crop is resistant to a wide range of diseases.

Mature Sugar Graze promotes good weight gains and

provides adequate energy for livestock (Pacific Seeds

2009). Jumbo Plus, a forage sorghum hybrid cultivar, has

excellent re-growth potential and high productivity and is

adapted to both dryland and irrigated situations. It has

similar CP concentration to Sugar Graze with 56‒64% dry

matter (DM) digestibility when the plant is 55‒60 days

old or at 5‒10% flowering stage and can be used for

grazing, silage making and rotational cropping (Forage

Sorghum Guide 2015).

These crops have the potential to compete favorably

with maize silage in terms of yield and nutritive value

(Ketterings et al. 2005) and may be an appropriate

alternative to maize for utilizing irrigation water in

drought-prone areas. The shortage of ground water is the

primary limitation to cultivating grass in the dry zone. As

such, it is essential to select a drought-tolerant grass/fodder

species, and Sri Lankan farmers cultivate fodder sorghum.

In an initial study 7 cuttings were achieved from a single

planting yielding 24 kg of fresh fodder/m2 from a single

cutting with plant spacing of 45 × 15 cm (Sivayoganathan

2016). While research on sorghum cultivars in Pakistan has

shown marked differences between cultivars in green

fodder yield and morphology under 30 cm row spacing

Figure 1. The experimental site, Kilinochchi District, in the

(Bakhsh et al. 2015), similar data on forage sorghum hybrid

dry zone of Sri Lanka.

Tropical Grasslands-Forrajes Tropicales (ISSN: 2346-3775)

36 H. Gnanagobal and J. Sinniah

The remaining period of the year is dry with the driest

done at weekly intervals. Plots were manually weeded at

period being June to August. During the study period

30 days after planting to reduce competition from weeds.

(August 2015‒January 2016), rainfall in individual

Five plants were randomly selected from each plot for

months varied greatly (Figure 2) and due to the extremely

recording of leaf length and width, leaf area [leaf area

low water retention capacity irrigation had to be applied

factor (0.72 for forage sorghum) × length × width] (Arkel

in some months.

1978), number of leaves per plant, stem girth, root length

According to Vavuniya meteorological data, the

from the base of the plant to tip of the selected average

average monthly temperature in the region is 28.4 °C

lengthier rootlet, internodal elongation and plant height at

(range 25.6‒30.0 °C), while maximum and minimum

weekly intervals. At 60 days after planting, on 3 October

averages are 35.0 and 21.3 °C, respectively (Jaffna and

2015, the crops were cut 15 cm from ground level and

Kilinochchi Water Supply and Sanitation Project 2010).

allowed to ratoon. Two ratoon cuts were made at 60-day

Soils of the area are red-yellow latosols (Haplustox),

intervals, on 1 December 2015 and 29 January 2016.

which are the most intensively cultivated soils of Jaffna

Immediately after harvesting, fresh forage was weighed

Peninsula and have very low inherent fertility. Extremely

using a spring balance. Samples (approximately 2 kg) of

poor water retention properties mean that dryland cropping

the harvested forage from each experimental plot were

is inappropriate, while conventional flood irrigation is

selected and air-dried for 24 hours, followed by oven-

impractical owing to very rapid infiltration and soil drying.

drying at 75 °C for 72 h to constant weight for dry matter

The experiment was laid out in a completely

yield determination. Data were subjected to Analysis of

randomized design (CRD) in a factorial arrangement of 3

Variance and mean separation was done with Duncan’s

plant spacings (30 × 15, 30 × 45 and 30 × 60 cm) and 2

multiple range test (P≤0.05) (Duncan 1955) with SPSS

cultivars (Sugar Graze and Jumbo Plus) with 3

(Statistical Package for Social Science) version 16.0 for

replications. Sowing was on 5 August 2015.

Windows.

Results

Initial harvest

For the initial harvest the 2 cultivars generally responded

differently to variation in plant spacing in terms of plant

morphology (Table 1).

While leaf length, leaf width and leaf area were

unaffected by plant spacing for Jumbo Plus, the narrow

spacing (30 × 15 cm) produced narrower leaves with

smaller area than the medium (30 × 45 cm) and wide (30

× 60 cm) spacings for Sugar Graze. Similarly, the narrow

Figure 2. Monthly rainfall of Kilinochchi District from

and wide spacings for Jumbo Plus produced more leaves

January 2015 to February 2016. Source: Department of Census

per plant than the medium spacing, while the medium

and Statistics of Sri Lanka (2015); Kilinochchi District (2016).

spacing for Sugar Graze produced more leaves than the

narrow spacing. In general, the thickest stems were

Within rows spacing was kept constant at 30 cm and

produced at the wide spacing and the thinnest at the

the spacing between rows was varied. Seeds were sown at

narrow spacing.

the rate of 2 seeds per hill and seedlings were thinned to

A similar trend occurred with root length with longer

a single plant per hill 2 weeks after sowing resulting in

root lengths generally being associated with wide plant

plant populations of approximately 222,000, 74,000 and

spacing and shorter root lengths with narrow spacing.

55,000 plants/ha for inter-row spacings of 15, 45 and 60

Plant height was unaffected by spacing for Jumbo Plus,

cm, respectively. Cattle manure was applied at planting at

while the wide spacing for Sugar Graze produced the

the rate of 100 kg/ha (N: 1.2‒1.9%, P: 0.2‒0.5%, K: 0.5‒

tallest plants.

1.1%) and inorganic fertilizers were applied 1 week after

Overall, Sugar Graze displayed slightly longer, wider

establishment of plants at the rate of 50 kg urea, 25 kg

leaves with much greater area than Jumbo Plus but

triple superphosphate and 12.5 kg muriate of potash/ha.

produced fewer leaves, though differences were not

During the dry spell of the study period, irrigation was

significant. This cultivar also produced longer roots than

Tropical Grasslands-Forrajes Tropicales (ISSN: 2346-3775)

Yield parameters of forage sorghum hybrids 37

Table 1. Effects of row spacing (30 x 15, 30 x 45, 30 x 60 cm) on growth parameters and yields of Sugar Graze and Jumbo Plus 60

days after planting (initial harvest).

Parameter

Sugar Graze

Jumbo Plus

30 × 15

30 × 45

30 × 60

30 × 15

30 × 45

30 × 60

Leaf length (cm)

90.6 ± 4.92a

91.4 ± 2.17a

92.8 ± 2.69a

86.5 ± 3.52a

89.5 ± 2.19a

87.6 ± 3.11a

Leaf width (cm)

8.6 ± 0.86b

9.2 ± 0.80a

9.2 ± 0.67a

7.27 ± 0.64a

7.37 ± 0.83a

7.15 ± 0.91a

Leaf area (cm2)

560 ± 64.5b

608 ± 61.1a

616 ± 48.8a

443 ± 36.6a

463 ± 66.7a

473 ± 67.9a

Number of leaves/plant

13.0 ± 0.93b

13.9 ± 0.83a

13.3 ± 0.96ab

15.0 ± 0.93a

13.7 ± 0.90b

15.1 ± 1.68a

Stem girth (cm)

5.50 ± 0.38b

6.23 ± 0.86a

6.65 ± 0.62a

5.65 ± 0.52b

6.03± 0.77b

6.57 ± 0.46a

Root length (cm)

23.5 ± 0.01c

28.3 ± 0.01a

26.0 ± 0.01b

22.3 ± 0.01c

24.3 ± 0.01b

25.6 ± 0.01a

Internodal elongation (cm)

21.9 ± 2.54a

20.6 ± 1.95a

21.4 ± 2.41a

20.8 ± 1.77a

22.7 ± 1.16a

23.2 ± 1.95a

Plant height (cm)

227 ± 22.7b

239 ± 40.6b

278 ± 18.5a

290 ± 14.8a

292 ± 14.3a

297 ± 25.4a

Dry matter yield (t/ha)

14.1 ± 2.60a

11.4 ± 1.94b

11.3 ± 1.80b

12.6 ± 2.77a

9.2 ± 2.81ab

6.3 ± 1.71b

Each value is a mean ± SD for 3 replicates.

Within rows and cultivars, means without a common letter differ (P≤0.05).

Jumbo Plus but internodal elongation was greater for

Dry matter yields followed similar trends in both

Jumbo Plus, resulting in taller plants.

cultivars with declining yields as plant spacing increased,

Dry matter (DM) yields were inversely related to plant

but differences were not always significant (Table 2).

spacing with yield decreasing progressively as plant

Varietal differences in DM yield were small.

spacing increased, although differences were not always

significant. Sugar Graze produced higher DM yields than

Second ratoon crop

Jumbo Plus (Table 1).

As for the first ratoon crop, row spacing had no significant

First ratoon crop

effect on leaf width, leaf area and stem girth in either

cultivar, while inconsistent responses occurred for the

In the first ratoon crop some parameters, viz. leaf length,

remaining morphological parameters (Table 3). There were

number of leaves and plant height, were not influenced by

consistent effects of row spacing on DM yields in both

spacing in either cultivar, while leaf width, leaf area,

cultivars with yields declining as plant spacing increased,

internodal elongation and stem girth varied inconsistently

but differences were significant (P<0.05) only for Jumbo

with row spacing in the 2 cultivars (Table 2). Varietal

Plus. Dry matter yields for Jumbo Plus at the medium and

differences also were noted among the morphological

wide spacings declined dramatically to about half those for

parameters of the first ratoon crop, where generally leaf length

Sugar Graze. Overall, increases in row spacing resulted in

and width, leaf area, number of leaves per plant and stem girth

greater percentage yield decreases in Jumbo Plus than in

were greater for Sugar Graze, while Jumbo Plus showed

Sugar Graze. Yields for both ratoon crops were generally

higher values for internodal elongation and plant height.

about 25‒35% of those obtained at the initial harvest.

Table 2. Effects of row spacing (30 x 15, 30 x 45, 30 x 60 cm) on growth parameters and yields of first ratoon crop of Sugar Graze and Jumbo Plus.

Parameter

Sugar Graze

Jumbo Plus

30 × 15

30 × 45

30 × 60

30 × 15

30 × 45

30 × 60

Leaf length (cm)

78.3 ± 5.07a

72.7 ± 20.2a

78.1 ± 3.93a

71.3 ± 4.78a

71.4 ± 5.28a

70.0 ± 5.49a

Leaf width (cm)

6.79 ± 0.70a

6.01 ± 0.66b

6.37 ± 0.52ab

4.19 ± 0.41b

4.57 ± 0.34a

4.45 ± 0.49ab

Leaf area (cm2)

382 ± 50.7a

317 ± 100b

358 ± 39.1ab

215 ± 29.4a

234 ± 22.8a

224 ± 35.6a

Number of leaves/plant

9.87 ± 0.83a

9.53 ± 1.25a

10.3 ± 1.03a

9.33 ± 1.18a

9.53 ± 1.13a

8.67 ± 1.50a

Stem girth (cm)

5.15 ± 0.36a

4.45 ± 0.34b

4.63 ± 0.45b

3.83 ± 0.37a

3.88 ± 0.15a

3.63 ± 0.21b

Internodal elongation (cm)

16.1 ± 4.07a

17.8 ± 2.62a

17.5 ± 2.17a

16.9 ± 3.32b

19.6 ± 2.16a

21.1 ± 1.18a

Plant height (cm)

115 ± 19.9a

115 ± 13.0a

119 ± 14.6a

131 ± 15.8a

138 ± 11.6a

139 ± 17.4a

Dry matter yield (t/ha)

3.36 ± 0.531a 2.95 ± 0.614a 2.36 ± 0.416a 3.84 ± 0.511a 3.27 ± 0.309a 2.13 ± 0.483b

Each value is a mean ± SD for 3 replicates.

Within rows and cultivars, means without a common letter differ (P≤0.05).

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38 H. Gnanagobal and J. Sinniah

Table 3. Effects of row spacing (30 x 15, 30 x 45, 30 x 60 cm) on growth parameters and yields of second ratoon crop of Sugar Graze and Jumbo Plus.

Parameter

Sugar Graze

Jumbo Plus

30 × 15

30 × 45

30 × 60

30 × 15

30 × 45

30 × 60

Leaf length (cm)

62.7 ± 6.15a

59.0 ± 3.58a

57.5 ± 13.1a

60.7 ± 5.57a

51.5 ± 7.15b

54.5 ± 4.63ab

Leaf width (cm)

5.78 ± 0.78a

5.73 ± 0.82a

5.65 ± 0.98a

3.63 ± 0.47a

3.68 ± 0.62a

4.13 ± 0.38a

Leaf area (cm2)

262 ± 50.7a

244 ± 45.4a

239 ± 89.0a

158 ± 18.5a

137 ± 35.0a

162 ± 21.7a

Number of leaves/plant

7.00 ± 0.63b

7.00 ± 0.01b

8.33 ± 0.52a

6.00 ± 0.63a

6.00 ± 0.63a

6.66 ± 0.82a

Stem girth (cm)

4.50 ± 0.55a

4.25 ± 0.42a

4.08 ± 0.38a

3.17 ± 0.26a

3.20 ± 0.32a

3.36 ± 0.22a

Internodal elongation (cm)

20.0 ± 2.83a

22.1 ± 2.13a

19.0 ± 1.95b

25.2 ± 3.95a

26.7 ± 3.36a

24.4 ± 2.48a

Plant height (cm)

190 ± 17.2a

178 ± 10.2ab

167 ± 10.1b

191 ± 9.12a

182 ± 8.91a

168 ± 9.31b

Dry matter yield (t/ha)

4.47± 0.744a 3.29 ± 1.090a 2.85 ± 0.350a 4.33 ± 0.358a 1.77 ± 0.206b 1.42 ± 0.539b

Each value is a mean ± SD for 3 replicates.

Within rows and cultivars, means without a common letter differ (P≤0.05).

Discussion

(460 ± 58.9 cm2) may be due to differences in genetic

makeup of the cultivars. Musa et al. (1993), Naeem et al.

Leaf length and width

(2002), Mahmud et al. (2003) and Chohan et al. (2003;

2006) also observed variation in leaf area among various

Leaf development has been described extensively for

cultivars and varieties of forage sorghum.

fodders, as growth is mostly reflected in large increases in

leaf length as plants grow to maturity, accompanied by

Number of leaves per plant

relatively small increases in width and thickness (Skinner

and Nelson 1994). Large leaf lengths are also important

The general absence of any consistent effect of row spacing

for the survival of individual plants within a sward (Barre

on leaf number per plant is in agreement with the findings

et al. 2015). Leaf length and width values observed for

of Liu et al. (2004), who observed for maize that it did not

both cultivars during the present study were slightly

affect leaf number. In contrast Lamana (2007) reported that

greater than the values recorded by Singh et al. (2014) for

wider plant spacing in maize had a positive effect on

leaf length (45‒70 cm) and width (4‒7 cm) of sorghum

number of leaves. The values recorded for number of

hybrids. Leaf length of Sugar Graze was similar to the 95

leaves per plant for both cultivars in the present study were

± 2.0 cm reported by Pahuja et al. (2014) in India, for the

consistent with those of Monteiro et al. (2012), who

first cut at 50% flowering and a spacing of 15 × 45 cm,

reported that number of leaves in forage sorghum is

whereas leaf width was slightly higher than that recorded

generally between 14 and 17. Chohan et al. (2003) and

by the same authors (6 ± 0.58 cm).

Naeem et al. (2002) also reported variation among different

cultivars of sorghum for number of leaves per plant.

Leaf area

Stem girth

The results of the present study demonstrate that leaf area

increases as plant spacing increases as shown by Lamana

Stem girth recorded in the present study was similar to

(2007). This could be due to less competition among

that reported by Pahuja et al. (2014) in India for stem girth

plants for space and soil nutrients as the plant population

of Sugar Graze (5.9 ± 0.21 cm at 50% flowering stage and

per unit area decreased. Therefore, the lower population

15 × 45 cm spacing). While Yosef et al. (2009) and Ayub

density which resulted from the wider plant spacing gives

et al. (1999) found significant variation in stem diameter

better conditions for more accumulation of photosynthetic

among different cultivars of sorghum, cultivar differences

products, better growth and expansion of foliage, which

in our study were not statistically significant (P>0.05).

was in turn expressed in greater DM yields. The range of

values for leaf area (440‒615 cm2) for Sugar Graze and

Root length

Jumbo Plus in the present study were in agreement with

the values reported by Nabi et al. (2006) for advanced

The trend for root length to increase as row spacing

lines of forage sorghum cultivars. The higher mean leaf

increased would reflect greater competition between

area in Sugar Graze (595 ± 62.3 cm2) than in Jumbo Plus

plants at the narrower spacings. Despite the shorter root

Tropical Grasslands-Forrajes Tropicales (ISSN: 2346-3775)

Yield parameters of forage sorghum hybrids 39

length per plant at the narrow spacing, the much greater

cultivars. Epasinghe et al . (2012) reported DM yields of

plant populations at this spacing would have resulted in

Sugar Graze in Sri Lanka of 5,230 kg/ha at 60 days after

substantially greater root length per unit area than at the

planting at 45 × 15 cm spacing and these lower yields

wider spacings. As a result plants at the narrow spacing

might be attributed to the differences in the spacing, soil

would have had better opportunity to utilize soil water and

fertility and environmental conditions. By contrast Nabi

nutrients than at wider spacing, resulting in higher DM

et al. (2006) recorded yields of 10,400‒13,100 kg DM/ha

yields.

for advanced lines of forage sorghums and Silungwe

(2011) recorded 13,262 kg DM/ha at 15 cm row spacing

Internodal elongation and plant height

78 days after sowing for Sugar Graze.

Forage yield is a function of growth parameters, viz.

Plant height as a growth parameter is a result of elongation

plant population, plant height, leaf:stem ratio, leaf area,

of the stem internodes, which is influenced by the

and leaf area index (Lamana 2007). The differences in

environment as suggested by Weston (1967). In the current

DM yield between the 2 cultivars could be attributed to

study taller plants were observed with wider spacing,

the fact that Sugar Graze exceeded Jumbo Plus in the

which contrasts with reports in the literature that narrower

growth parameters leaf length and width, leaf area and

spacing will give taller plants as a result of competition for

root length. Watson (1947) has shown that variation in

sunlight (Lamana 2007). However, the absence of any

total dry weight of plants is more dependent on variation

effect of plant spacing on plant height of Jumbo Plus

in leaf area. Light interception capacity of the leaf is

supports the finding of Roy and Biswas (1992) that plant

amplified with the increase in leaf area often leading to

height at maturity was not affected by plant spacing. The

increase in photosynthesis and DM yield. Therefore,

significant differences in plant height between the 2

higher DM yield recorded for Sugar Graze might be

cultivars may be due to genotypic variation, as differences

attributed to its higher leaf area than Jumbo Plus.

in internodal elongation between varieties can lead to

differences in height as reported by Evans (1975) and

First and second ratoon crops

Weston (1967). Nabi et al. (2006) and Silungwe (2011),

who worked with different forage sorghum cultivars, also

The most consistent findings with the ratoon crops were

reported plant heights (203‒230 cm) lower than those in

that there were fewer leaves per plant, leaves were

the present study (227‒298 cm), as did Pahuja et al. (2014)

smaller, height was less and DM yields were lower than

for Sugar Graze (189 ± 1.9 cm) in India.

for the initial harvest. However, DM yield remained a

factor of plant spacing with higher yields at narrower row

Dry matter yield

spacing, indicating that plants were still accessing

moisture and nutrients from the soil in sufficient

Plant spacing has a marked impact on the efficiency of

quantities to maintain acceptable growth levels. The

use of land, light, water and nutrients. By optimizing plant

reduced yields are possibly a function of nutrient supply

spacing, highest yield potential can be achieved from the

in the soil being depleted by the initial crop and a change

smallest possible area (Oseni and Fawusi 1986). The

in seasonal conditions over time. There were no

direct relationship between DM yield and plant

significant differences between Sugar Graze and Jumbo

population agrees with the findings of Fisher and Wilson

Plus in DM yields for the first ratoon crop, in contrast with

(1975), who reported greater DM yield with higher plant

the generally higher yields for Sugar Graze in the initial

populations than with lower plant populations. Wolf et al .

crop and second ratoon crop. Despite having smaller

(1993) and Graybill et al. (1991) also reported that DM

leaves and thinner stems than Sugar Graze, the greater

yield of forage maize responded positively to plant

height of Jumbo Plus ensured that yields in the 2 cultivars

density. This relationship would be affected by the

were similar. The success of the second crop is often a

availability of soil moisture, and the application of

function of how early the main crop was planted and

irrigation on a regular basis in this study would have

harvested, which determines the seasonal conditions

ensured that all row spacings/plant populations had

under which the first and second ratoon crops must grow.

adequate water. Dry matter yield recorded for Sugar

However, normally ratoon crops of sorghum are expected

Graze in the current study seemed to be less affected by

to yield from 25 to 35% of the main crop (Livingston and

differences in row spacing than Jumbo Plus, which

Coffman 1996), and our yields fall within this range.

appeared not to be related to root length as there were no

Significant differences in DM yield between main and

significant differences in root length between the

ratoon crops have been reported by Saberi and Aishah

Tropical Grasslands-Forrajes Tropicales (ISSN: 2346-3775)

40 H. Gnanagobal and J. Sinniah

(2014), when assessing yield responses of forage ratoon

Duncan DB. 1955. Multiple range and multiple F tests.

sorghum under varying salinity levels and irrigation

Biometrics 11:1–42. DOI: 10.2307/3001478

frequencies.

Epasinghe TM; Jayawardena VP; Premalal GGC. 2012.

Our findings suggest that both Sugar Graze and Jumbo

Comparison of growth, yield and nutritive value of maize,

multi-cut fodder sorghum and hybrid Napier (var. Co3) grown

Plus will grow satisfactorily under irrigation in this

in wet zone of Sri Lanka. In: Kodithuwakku SP; Himali SMC,

environment. While DM yields from the first harvest were

eds. Proceedings of 22nd Annual Students Research Session.

excellent, yields from the ratoon crops were significantly

Department of Animal Science, University of Peradeniya, Sri

lower despite the application of irrigation. A plant spacing

Lanka, 30 November 2012. p. 23. goo.gl/Ri3AFR

of 30 × 15 cm produced the highest yields but results

Evans LT. 1975. The physiological basis of crop yield. In:

under rain-fed conditions would not necessarily be the

Evans LT, ed. Crop physiology, some case histories.

same. Further studies to determine the performance in the

Cambridge University Press, Cambridge, UK. p. 327–335.

low-rainfall ( yala) season are necessary to determine

Fisher KS; Wilson GL. 1975. Studies of grain production in

Sorghum bicolor (L.) Moench. V. Effect of planting density

desirable spacings under such dry conditions. Chemical

on growth and yield. Australian Journal of Agricultural

analyses of forage and digestion studies would provide

Research 26:31–41. DOI: 10.1071/ar9750031

valuable information on the relative merits of these two

Forage Sorghum Guide. 2015. goo.gl/ALg81T (accessed 25

cultivars for livestock feeding.

December 2016).

Graybill JS; Cox WJ; Olis DJ. 1991. Yield and quality of forage

References

maize as influenced by hybrid and planting date and plant

density. Agronomy Journal 83:559–564. DOI: 10.2134/

Arkel HV van. 1978. Leaf area determinations in sorghum and

agronj1991.00021962008300030008x

maize by the length-width method. Netherlands Journal of

Jaffna and Kilinochchi Water Supply and Sanitation Project.

Agricultural Science 26:170–180.

2010. Jaffna and Kilinochchi Water Supply and Sanitation

Ayub M; Tanveer A; Mahmud K; Ali A; Azam M. 1999. Effect

Project: Initial environmental examination. Asian Develop-

of nitrogen and phosphorus on the fodder yield and quality

ment Bank, Metro Manila, Philippines. goo.gl/jCkAtK

of two sorghum cultivars ( Sorghum bicolor L.). Pakistan

Ketterings QM; Godwin G; Cherney JH; Kilcer TF. 2005.

Journal of Biological Science 2:247–250. DOI: 10.3923/

Potassium management for brown midrib sorghum x

pjbs.1999.247.250

sudangrass as replacement for corn silage in the North-eastern

Bakhsh A; Zahid MS; Shafeeq S; Gurmani ZA; Khan S. 2015.

USA. Journal of Agronomy and Crop Science 191:41–46.

Comparison of green fodder yield in sorghum varieties

DOI: 10.1111/j.1439-037x.2004.00144.x

under rainfed conditions of Islamabad. Life Sciences

Kilinochchi District. 2016. Rainfall pattern of district.

International Journal 9:3146–3149. goo.gl/GUCEdg

Performance report and annual accounts - 2016. Kilinochchi,

Barre P; Turner LB; Escobar-Gutiérrez AJ. 2015. Leaf length

NP, Sri Lanka. p. 12. goo.gl/SS6fUx

variation in perennial forage grasses (Review). Agriculture

Lamana MCL. 2007. Effect of spacing between plants on

5:682–696. DOI: 10.3390/agriculture5030682

growth and forage yield of two maize ( Zea mays L. )

Bilal MQ; Abdullah M; Lateef M. 2001. Effect of Mott dwarf

cultivars. M.Sc. Thesis. University of Khartoum, Khartoum,

elephant grass ( Pennisetum purpureum) silage on dry matter

Sudan. goo.gl/fGnqfx

intake,

milk

production,

digestibility

and

rumen

Liu W; Tollenaar M; Stewart G; Deen W. 2004. Within-row plant

characteristics in Nili-Ravi buffaloes. Proceedings of 54th

spacing variability does not affect corn yield. Agronomy

Annual Reciprocal Meat Conference, Vol. II, Indianapolis,

Journal 96:275–280.goo.gl/HgDKYZ

IN, USA. p. 24–28.

Livingston SD; Coffman CG. 1996. Ratooning grain sorghum

Chohan MSM; Naeem M; Khan AH; Salah-ud-Din S. 2003.

on the Texas Gulf Coast. Texas Agricultural Extension

Performance of newly developed forage varieties of

Service. The Texas A&M University System, College

sorghum ( Sorghum bicolor (L.) Moench). Asian Journal of

Station, TX, USA. hdl.handle.net/1969.1/87772

Plant Sciences 2:48–50. DOI: 10.3923/ajps.2003.48.50

Mahmud K; Ahmad I; Ayub M. 2003. Effect of nitrogen and

Chohan MSM; Naeem M; Khan AH; Kainth RA. 2006.

phosphorus on the fodder yield and quality of two sorghum

Performance of pearl millet ( Pennisetum americanum)

( Sorghum bicolor L.) cultivars. International Journal of

varieties for green fodder yield. Journal of Agricultural

Agricultural Biology 5:61–63. goo.gl/5kQMRk

Research 44:23–27. goo.gl/BqYrDJ

Monteiro JST; Havrland B; Ivanova T. 2012. Sweet Sorghum

Department of Census and Statistics of Sri Lanka. 2015.

( Sorghum bicolor (L.) Moench) bioenergy value – Importance

Monthly rainfall in district (Kilinochchi) 2012–2015.

for Portugal. Agricultura Tropica et Subtropica 45:12–19. DOI:

goo.gl/xeAUhj

10.2478/v10295-012-0002-y

Douglas J. 1980. Yield of crops for forage and fodder. In: Drew

Musa M; Gondal MR; Ali A; Ashraf M. 1993. Effect of cutting

KR; Fennessy PF, eds. Supplementary feeding. Occasional

at different plant growth stages and heights above ground

Publication No. 7. New Zealand Society of Animal Production,

levels on green fodder and dry matter yields of Sadabahar.

Mosgiel, New Zealand. p. 1–47.

Journal of Agricultural Research 31:409–413.

Tropical Grasslands-Forrajes Tropicales (ISSN: 2346-3775)

Yield parameters of forage sorghum hybrids 41

Nabi CG; Riaz M; Ahmad G. 2006. Comparison of some

M.Sc. Thesis. Massey University, Palmerston North, New

advanced lines of Sorghum bicolor (L.) Moench for green

Zealand. hdl.handle.net/10179/3234

fodder/dry matter yield and morpho-economic parameters.

Singh PK; Gautam RK; Roy SD; Singh AK; Ahmed SKZ. 2014.

Journal of Agricultural Research 44:191–196. goo.gl/imqKrC

Performance of sorghum varieties for fodder yield during

Naeem M; Chauhan MSM; Khan AH; Salahaddin S. 2002.

Rabi season in Andaman & Nicobar Islands. Journal of the

Evaluation of different varieties of sorghum for green fodder

Andaman Science Association 19:174–176. goo.gl/Db94KU

yield potential. Asian Journal of Plant Sciences 1:142–143.

Sivayoganathan B. 2016. The SOLID approach to producing

DOI: 10.3923/ajps.2002.142.143

year-round cattle feed. United States Agency for International

Oseni TO; Fawusi MO. 1986. Influence of nursery spacing and

Development (USAID), Colombo, Sri Lanka. goo.gl/

plant arrangement on growth and leaf nutrient content of

k1wvpW

three citrus root stock seedlings. Tropical Agriculture

Skinner RH; Nelson CJ. 1994. Role of leaf appearance rate and

64:41‒45. goo.gl/g5rVQW

the coleoptile tiller in regulating tiller production. Crop

Pacific Seeds. 2009. Sugar Graze for grazing, winter stand over

Science 34:71–75. DOI: 10.2135/cropsci1994.0011183x003

and pit silage. goo.gl/zMvyLV (accessed 25 December 2015).

400010013x

Pahuja S; Arya S; Kumari S; Panchta R. 2014. Evaluation of

Watson D. 1947. Comparative physiological studies on the growth

forage sorghum hybrids [ Sorghum bicolor (L.) Moench].

of field crops. II. The effect of varying nutrient supply on net

Forage Research 40:159–162. goo.gl/4BCD3W

assimilation rate and leaf area. Annals of Botany 11:375‒407.

Roy SK; Biswas PK. 1992. Effect of plant density and

jstor.org/stable/42907023

detopping following silking on cob growth, fodder and grain

Weston RH. 1967. Factors limiting the intake of feed by sheep. II.

yield of maize ( Zea mays). The Journal of Agricultural

Studies with wheaten hay. Australian Journal of Agricultural

Science 119:297–301. DOI: 10.1017/s0021859600012156

Research 18:983–1002. DOI: 10.1071/ar9670983

Saberi AR; Aishah HS. 2014. Physiological effects on re-

Wolf DP; Coors TG; Albrecht KA; Undersonder DJ; Carter PR.

growth of forage sorghums ratoon crop under varying

1993. Agronomic evaluations of maize genotypes selected for

salinity and irrigation frequency. British Journal of Applied

extreme fibre concentration. Crop Science 33:1359–1365.

Science & Technology 4:2277–2289. DOI: 10.9734/bjast/

DOI: 10.2135/cropsci1993.0011183x003300060047x

2014/5738

Yosef E; Carmi A; Nikbachat M; Zenou A; Umiel N; Miron J.

Sarwar M; Khan MA; Iqbal Z. 2002. Feed resources for

2009. Characteristics of tall versus short-type varieties of

livestock in Pakistan. International Journal of Agriculture &

forage sorghum grown under two irrigation levels, for

Biology 4:186–192. goo.gl/X2TVR2

summer and subsequent fall harvests, and digestibility by

Silungwe D. 2011. Evaluation of forage yield and quality of

sheep of their silages. Animal Feed Science and Technology

sorghum, sudangrass and pearl millet cultivars in Manawatu.

152:1–11. DOI: 10.1016/j.anifeedsci.2009.01.018

(Received for publication 14 February 2017; accepted 29 September 2017; published 31 January 2018)

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Tropical Grasslands-Forrajes Tropicales (ISSN: 2346-3775)